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The story and concept of the episode was written by series co-creator Chuck Lorre. Upon airing, "The Recombination Hypothesis" garnered The episode begins with Sheldon receiving a cardboard cut-out of Spock he ordered, however is dismayed to find it is not the Spock played by Leonard Nimoy but rather Zachary Quinto. While Sheldon complains, Leonard sees Penny and they exchange smiles. Leonard walks over asking her if she has any plans for dinner.
Penny looks surprised about being asked out on a date and accepts. As Penny nervously tries on clothes with Amy and Bernadette, Bernadette assures her that everything will work out since Leonard is crazy about her. Whilst playing Settlers of CatanSheldon, Howard and Raj ask Leonard why he is putting himself through another relationship with Penny, as they had broken up two years ago when things got too serious for her.
During dinner, Leonard and Penny find their first date in two years awkward, and decide to make-believe they are on a real first date. They both tell each other about their work and Penny is amused when Leonard boasts that he built a Big bang penny and leonard start dating and calls himself "King of the Nerds".
Leonard then asks Penny whether they are going to get back together. A shocked Penny confesses that she will always have feelings for him and that he always overthinks everything. Leonard is offended by this and returns home dejected. Later, Leonard is asleep and awakened by a text from Penny who meets him in the hallway, kisses him, takes him into her apartment and tells him to be quiet. Leonard wonders how a bad date ended in sex. Penny thought that things were great until he asked if they were getting back together. Leonard suggests that they try being together and not tell their friends until they know it is working, treating their relationship like a new version of software, a "Penny and Leonard 2.
At The Cheesecake Factory, the group discuss how the date did not work out. Leonard sits down and starts an argument with Penny by insulting her.
Later that night, Penny demands to know why Leonard was a jerk at work since they had agreed to be friends, not fight. Leonard he toward her apartment because their new relationship is about sex after fighting. Later, Leonard asks Penny what exactly they are doing since every relationship scenario he plays out ends badly. When Penny reminds him that he overthinks everything, Leonard hears Sheldon calling him. It turns out that Leonard was daydreaming, and he is still with Sheldon complaining about the Spock cardboard cut-out. Leonard decides that, despite the fact it seems that the relationship will once again end badly, he still wishes to ask Penny out.
Penny asks whether he has thought things through. Leonard replies that he has and still thinks they should go ahead Big bang penny and leonard start dating it. Penny therefore agrees and then smiles. In Penny's bedroom, she is again choosing an outfit for her date. Daydreaming, she sees herself heavily pregnant during a wedding ceremony with Leonard. She snaps out of the daydream and decides to go to the drugstore before the date. The first scene is a salute to the original pilot where Leonard and Sheldon climb the stairs wearing similar outfits and they spot Penny through her apartment door wearing the same shirt from the pilot.
L Shaffer of IGN gave the episode a "really good" review. Day noted that revisiting Leonard and Penny on the th episode was the best way to honor the history of the series. Though she was happy to see "Peonard or Lenny" back together, she felt that the other guys had the best lines with the "wood" jokes. Overall she found the show "brought us back to the origins of the comedy and what made us fall in love with these nerds and their gorgeous neighbor.
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